Genus Tridentoinfundibulum
Tshoijo, in Popova, 1958
General. Middle-sized Cyathostominae. MC flattened, divided
into inner and outer rings. Posterior edge of MC posterior to edge of BC. Amphids
not markedly projected through MC surface. Tip and longer stalk of submedian papillae
extend through MC. Tip of submedian papillae bulbous, 2 times long as thick. Stalk
of submedian papillae broader than long. Elements of ELC longer than broad, tips
pointed; insertion point on tips of ILC. Elements of ILC difficult to distinguish,
slightly longer than broad, identical in number to ELC; insertion point at 1/4
or less of BC depth. Line formed by insertion of elements of ILC straight. Form
of posterior edge of elements of ILC straight, unadorned. Support for ELC continuous
with BC, elongate, curving, thin at one end. Septum intracoronare origin on support.
Medial insertion of septum intracoronare situated anteriorly to junction of ELC
and ILC. Walls of BC straight, thicker in middle. Buccal cavity wider than deep,
cylindrical. Dorsal gutter short, broad at base, less than 1/2 of BC depth. Buccal
teeth absent. Esophageal funnel enlarged, especially in dorsal sector which forms
invagination containing 3 esophageal teeth . Anterior muscular portion of esophagus
about 1/4 to 1/3 of esophagus length. Excretory pore posterior to NR. Anterior
deirids near middle of glandular esophagus.
Male: Dorsal ray with 6 branches. Ventral rays shorter
than laterals. Dorsal lobe longer then lateral lobes. Externodorsal rays origin
at junction of dorsal and laterals rays. Gubernaculum large, with dorsal handle
and ventral notch. Genital cone short, conical. Spicule tips hook- or harpoon-shaped
Female: Vulva more than 1 tail length from anus. Vagina
shorter than sphincter of ovejector. Ovejector vestibule oval or Y-shaped, infundibulum
longer than sphincter. Tail conical, short, length less than 2x diameter at anus.
Type species: T. gobi Tshoijo
in Popova, 1958
Discussion
Although much remains to be learned about the generic relationships
of this species, only recently described in enough detail to be recognized (Dvojnos
and Kharchenko, 1994). this is a good species that has been collected in Asia,
North America and Europe (Lichtenfels et al., 1998). Lichtenfels and Klei (1988)
described this species as Cylicostephanus torbertiae. Tridentoinfundibullum is
recognized at genus level here because it appears on the basis of morphological
(Lichtenfels and Hoberg, unpublished) and molecular phylogenies (McDonnell et
al., 2000; Hung et al. 2000) to be within a clade that includes Cylicostephanus
and closer to, but separated from, Cylicodontophorus. Hartwich (1986) did not
study this species. |